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Janeane Garofalo plays Dr. Abby Barnes, the “Truth About Cats and Dogs” radio question-and-answer show host who unwittingly entices a listener over the radio with her soothing voice and personality. This listener, Brian, tries to meet the Abby from the radio, but Noelle, played by Uma Thurman, is mistaken for the real thing when Brian comes to the studio. Instead of clearing things up right away, the self-conscious Abby allows her best friend, Noelle, a tall, stunning blonde, to take her place for a while. Abby takes on the made-up persona of Donna, while thinking Brian would never go for her, a short, cute, brunette, who thinks she’s unattractive. As the real Abby woos Brian over the phone and radio, Noelle, the pseudo-Abby, takes her place in the flesh. As time goes on, Abby feels more and more confident that Brian would rather have the beautiful Noelle than the simply attractive Abby.

After their first year at college, the guys reunite for another summer of fun. Jim continues his quest for sexual independence by seeking the help of his old prom date, Michele, after an unexpected call from Nadia who plans to visit Jim. Meanwhile, Kev and Vicky find themselves in an awkward situation after having broken up for a year. Oz must deal with a long distance relationship when Heather heads off to France to study abroad. Old feuds die hard as the ever-so-horny Stifler harbors his hatred toward Finch, who is practicing the Hindu practice of Tantra.

Barry Egan (Adam Sandler) is a small business owner with seven sisters whose abuse has kept him alone and unable to fall in love. When a harmonium and a mysterious woman (Emily Watson) enter his life, his romantic journey begins.