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Marlin (a clown fish) is a widower who only has his son Nemo left of his family after a predator attack. Years later, on Nemo’s first day of school, he’s captured by a scuba diver and taken to live in a dentist office’s fish tank. Marlin and his new absent-minded friend Dory set off across the ocean to find Nemo, while Nemo and his tankmates scheme on how to get out of the tank before he becomes the dentist’s niece’s new pet.

Steve Zissou, sea-film auteur a la Jacques Cousteau, has reason to be melancholy: his partner has been eaten, perhaps by a mythic jaguar shark, his wife may be taking up with her ex-husband, a young man appears claiming Steve is his father (Steve hates fathers), his most recent films have tanked, he’s having trouble raising money for his venture to revenge his partner, and he’s attracted to a pregnant reporter who prefers the pretender. At sea, in pursuit of the shark, will he escape pirates and mutiny, forge the bonds of fatherhood, place his arm around his wife, find the monster of the deep, re-establish box office hegemony, and discover a reason to smile?

The sea underworld is shaken up when the son of the shark mob boss is found dead and a young fish named Oscar is found at the scene. Being a bottom feeder, Oscar takes advantage of the situation and makes himself look like he killed the finned mobster. Oscar soon comes to realize that his claim may have serious consequences.