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Numerous New York City-dwellers come to the exclusive club Shortbus to work out problems in their sexual relationships. Rob and Sophia are a happily married couple, except for the fact that she has never experienced sexual climax. This irony follows her to work, because she is a couples counselor who frequently has to deal with the sexual issues other couples have. Two of her patients are Jamie and James, a gay couple who have been monogamous for five years and counting. James wants to bring other men in to the relationship, and his own history with depression may hint at an ulterior motive. Ceth (Pronounced like Seth) may be the perfect addition to their family, but Caleb, a voyeur from across the street, may have his own ideas about that. Sophia visits Severin, a dominatrix with secrets of her own to reveal.

One week before her sex-change operation, Bree receives a call from a 17-year-old identifying himself as her son from a college liaison. Bree’s psychiatrist won’t approve the surgery until Bree deals with this relationship, so Bree flies to New York City, bails the youth out of juvenile detention, and offers him a ride back to Los Angeles without disclosing that she is his father. Both her plans and his go awry, and as secrets will out, what might become a friendship (or more) founders. The lad’s step-father, a sex-change support group, a peyote eater, a Navajo wrangler, and Bree’s family all play their parts in this exploration of family, gender, and expectations.