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A shipping disaster in the 19th Century has stranded a man and woman in the wilds of Africa. The lady is pregnant, and gives birth to a son in their tree house. Soon after, a family of apes stumble across the house and in the ensuing panic, both parents are killed. A female ape takes the tiny boy as a replacement for her own dead infant, and raises him as her son. Twenty years later, Captaine Phillippe D’Arnot discovers the man who thinks he is an ape. Evidence in the tree house leads him to believe that he is the direct descendant of the Earl of Greystoke, and thus takes it upon himself to return the man to civilization.

A young man lingers in the family home of his fiancee, after her accidental death. While grieving along with her parents and drawn into legal issues presented by a district attorney seeking justice for the family, he finds himself falling in love with another woman, against his own best intentions.

In 1936, the expatriated young and naive just-graduated British John Truscott (Hugh Dancy) arrives to the Sarawak, a British colony, to work in the Iban society. The beautiful Selima (Jessica Alba) is assigned to be his “sleeping dictionary”, to live and sleep with him and teach him the language and habits of the locals. The reluctant John and Selima fall in love for each other in a forbidden romance.